Regulation of the 11beta-hydroxysteroid dehydrogenasetype 2 promoter by steroid hormones in breast cancer cells. convergence of progesterone receptor binding to dna and jak/stat pathway activation

Tesis doctoral de Alicia Subtil Rodríguez

Steroid hormone receptors regulate gene expression interacting with target dna sequences but also activating cytoplasmic signalling pathways. Using the human 11beta-hydroxysteroid dehydrogenase type 2 (11 -hsd2) gene as a model, we investigated the contribution of both effects on a human progesterone responsive promoter in breast cancer cells. Our results suggest two different mechanisms of hormone-induced pr recruitment to the 11 -hsd2 promoter: i,) direct pr binding to dna at the proximal promoter, abolished when pr contains a mutated dna binding domain (dbd); and ii,) stat5a-mediated recruitment of pr to the distal promoter region, impaired by a jak/stat pathway inhibitor. Therefore, coordination of pr transcriptional effects and cytoplasmic signalling activation, in particular jak/stat pathway, are critical to regulate progestins-induced endogenous 11 -hsd2 gene expression in breast cancer cells. This is not unique to this promoter, as the jak/stat inhibitor also alters expression of other progesterone-regulated genes. Additionally, we report rna polymerase ii tracking from the distal region upon pr and stat5a binding and we propose that this region works as hormone-dependent transcriptional enhancer.

 

Datos académicos de la tesis doctoral «Regulation of the 11beta-hydroxysteroid dehydrogenasetype 2 promoter by steroid hormones in breast cancer cells. convergence of progesterone receptor binding to dna and jak/stat pathway activation«

  • Título de la tesis:  Regulation of the 11beta-hydroxysteroid dehydrogenasetype 2 promoter by steroid hormones in breast cancer cells. convergence of progesterone receptor binding to dna and jak/stat pathway activation
  • Autor:  Alicia Subtil Rodríguez
  • Universidad:  Pompeu fabra
  • Fecha de lectura de la tesis:  22/06/2007

 

Dirección y tribunal

  • Director de la tesis
    • Albert Jordan Vallés
  • Tribunal
    • Presidente del tribunal: ana Aranda iriarte
    • jordi Bernués martínez (vocal)
    • luciano Di croce (vocal)
    • Fernández de mattos sílvia (vocal)

 

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